The story begins with a strange black tower of energy, which would later come to be known as the "Pillar of Darkness", appearing in the Sea of Japan off the coast of Sadogashima. The tower's appearance causes seismic fluctuations that confound geologist Joe Hayakawa and his team. Returning home, Joe finds his pregnant wife Marie outside setting up equipment with which to study the phenomenon. Suddenly, a large crimson dragon-like creature with what looks like a gun barrel for a head emerges from the Pillar, and proceeds to lay to waste the military forces that try to attack it. When all seems lost, another dragon appears from the Pillar; this one silver-white and serpentine in appearance, with what looks like a blade for a head. The two creatures proceed to destroy each other, causing a reaction in the Pillar of Darkness.
Twelve years later, fraternal twins Ai and Yu Hayakawa find and board an interdimensional train to Wonderland, a strange world connected to Earth by the Pillar of Darkness, to find their parents, Joe and Marie, who have not returned from their latest of many trips there. On the train they meet Lisa Pacifist, a woman with special powers known as Kigen Arts.
When the trio arrive in Wonderland, they find out just how different it is from Earth, meeting a number of allies and enemies. Allies include Chobi, a chocobo who quickly takes a mutual liking to Yu and joins the group on their journey, and Kaze, a mysterious amnesiac gunman with a grudge against a sword-wielding warrior clad in white who he believes was responsible for the destruction of his homeworld. Kaze wields the Magun, a gun powered by bullets filled with "Soil" which summons a variety of great creatures to destroy his enemies.
Enemies include Earl Tyrant, a spoiled brat of a child living in the floating fortress palace of Gaudium as well as the ruler of Wonderland. Tyrant has four underlings referred to collectively as the Lords of Gaudium: Fungus, a humanoid mushroom man with super strength and regenerative powers; Herba, a humanoid plant woman with a shallow, self-centred attitude; Pist Shaz XI, a humanoid shark creature made of water who takes pride in analyzing the data of his opponents; and Oscha, a mysterious black humanoid figure whose face is always hidden behind a mask and Tyrant's closest servant. Also in service of the Earl is Makenshi, a sword-wielding warrior clad in white, though he prefers to be alone, staying away from the antics in the throne room, and seems to have his own agenda.
Lisa and the twins also find allies in the Comodeen, a rebel army group seeking to overthrow the Earl and establish true and fair order in Wonderland. Led by Knave, a strong man with a romantic nature, the Comodeen is made up entirely of volunteer members whose homeworlds were destroyed by Chaos. Despite the strength of their opponents, the Comodeen are a force to be reckoned with thanks to battle plans formulated by Miles, a tomboyish woman and second-in-command, and equipment and other supplies created by Cid, the resident mechanical mastermind who feels a little too strongly for his creations.
Early on in their journey, the group are accosted by a part of Omega, a mysterious being that destroys everything in its path simply by existing. Omega was defeated at some point prior to the beginning of the series, its body shattered into component pieces and forced into dormancy, during which it takes the form of a translucent pink crystal, referred to as an Omega Crystal. Though many pieces are still dormant during the series, three have awakened and attack the Interdimensional Train. As explained by Lou, a lycanthrope girl the group meet on the train who travels with the group for a while, developing a major crush on Kaze, once a part of Omega awakens, it constantly searches for its "siblings", other Omega Crystals, to eventually reform into the full creature. Eventually, one of the parts of Omega succeeds in destroying the Train, revealing the reason it was attacking it: the Interdimensional Train is powered by an Omega Crystal.
At one point, the group stops in an endless ....STREAM -------------------------------------------
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